Kabwe Cranium, or Broken Hill 1
"Rhodesian Man"
| | | Species: | Homo heidelbergensis | | Age: | 125,000 to 300,000 years | | Date of Discovery: | June 17, 1921 | | Location: | Kabwe, Zambia | | Discovered by: | Tom Zwiglaar | |
This cranium was the first early human fossil to be found in Africa. Miners looking for metal deposits in limestone caves discovered it. The find was later sent to Arthur Smith Woodward, who described the fossil in an article in volume 108 of the journal Nature, and gave it a new species name "Homo rhodesiensis" (Rhodesian Man). Once thought to be less than 40,000 years old, the Kabwe skull (also known as the Broken Hill skull) was used at one time to validate the supposed "primitiveness" of African peoples, demonstrating that while Europeans had evolved to the "level" of Cro-Magnon, African populations still looked essentially like Homo erectus. This assumption was shown to be flawed on many accounts, most crucially in that the date for this site based on the associated animal fossils found is at least 125,000 years old, and is probably significantly older. Some researchers have proposed that Kabwe may be a member of the African population from which all modern humans descended, although this cannot be definitively proven. The braincase profile is low and slopes back from a large supraorbital torus reminiscent of earlier H. erectus specimens. There is also the remnant of a sagittal keel and an occipital torus at the back of the skull, also recalling H. erectus. However, the face is more modern in appearance (less prognathic, flatter) and the brain size of about 1300 cc. is larger than seen in H. erectus. Thus, this cranium preserves many traits that are reminiscent of earlier H. erectus and hints of more modern traits known later in H. sapiens. The cranium shows evidence of disease and wounds that occurred in the lifetime of this individual. Ten of the upper teeth have cavities, and dental abscesses of the upper jaw are clearly visible in the upper photograph (above the right incisor/canine) and the middle photograph (above the first molar). Additionally, a partially healed wound is visible in the bottom two photographs, above and anterior of the hole for the ear. This wound measured roughly a quarter-inch across, and was made by either a piercing instrument or the tooth of a carnivore. Exactly which is unclear
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The lower inner surface of the neurocranium
The skull also contains the
sinus cavities, which are air-filled cavities lined with
respiratory epithelium, which also lines the large airways. The exact functions of the sinuses are debatable; they contribute to lessening the weight of the skull with a minimal reduction in strength, they contribute to resonance of the voice, and assist in the warming and moistening of air drawn in through the nasal cavities.
Development of the skull
The skull is a complex structure; its bones are formed both by
intramembranous and
endochondral ossification. The bones of the splanchnocranium (face) and the sides and roof of the neurocranium are formed by intramembranous (or
dermal) ossification, while the bones supporting the brain (the
occipital,
sphenoid,
temporal, and
ethmoid) are largely formed by endochondral ossification.
At birth, the human skull is made up of 404 separate bony elements. As growth occurs, many of these bony elements gradually fuse together into solid bone (for example, the
frontal bone). The bones of the roof of the skull are initially separated by regions of dense
connective tissue called "
cranial sutures". There are five sutures: the
frontal suture,
sagittal suture,
lambdoid suture,
coronal suture, and
squamosal suture. At birth these regions are fibrous and moveable, necessary for birth and later growth. This growth can put a large amount of tension on the "obstetrical hinge," which is where the squamous and lateral parts of the occipital bone meet. A possible complication of this tension is rupture of the
great cerebral vein of Galen. Larger regions of connective tissue where multiple sutures meet are called
fontanelles. The six fontanelles are: the anterior fontanelle, the posterior fontanelle, the two sphenoid fontanelles, and the two mastoid fontanelles. As growth and ossification progress, the connective tissue of the fontanelles is invaded and replaced by bone. The posterior fontanelle usually closes by eight weeks, but the anterior fontanelle can remain open up to eighteen months. The anterior fontanelle is located at the junction of the frontal and parietal bones; it is a "soft spot" on a baby's forehead. Careful observation will show that you can count a baby's heart rate by observing his or her pulse pulsing softly through the anterior fontanelle.
PathologyIf the brain is bruised or injured it can be life-threatening. Normally the skull protects the brain from damage through its hard unyieldingness, the skull is one of the most durable substances found in nature. In some cases, however, of
head injury, there can be raised
intracranial pressure through mechanisms such as a
subdural haematoma. In these cases the raised
intracranial pressure can cause herniation of the brain out of the
foramen magnum ('coning') because there is no space for the brain to expand; this can result in significant
brain damage or death unless an urgent operation is performed to relieve the pressure. This is why patients with
concussion must be watched extremely carefully.